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Echinoderm

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Crown-of-Thorns StarfishCrown-of-Thorns Starfish
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I

Introduction

Echinoderm, common name for about 6,000 living species constituting a phylum of marine animals, such as starfish, brittle stars, sea urchins, sand dollars, and sea cucumbers. They usually show a superficial five-part radial symmetry, and are generally equipped with peculiar tube feet. The phylum name is derived from the spiny skin.

The larval stages of this distinct and highly modified group are rather like those of the acorn worm, suggesting a relationship to the chordates. In later development, however, the left side of the body grows at the expense of the right, and the adults are quite unlike any other animals.

II

Structure

An echinoderm such as the starfish typically has a centrally positioned mouth surrounded by five arms that bear minute, fleshy tube feet with which the animal clings and crawls. The tube feet are supported by a complex internal hydraulic system that is inflated with sea water. Some echinoderms, especially brittle stars, crawl or swim by moving their arms. Often, as in sea urchins, rows of tube feet extend along the body surface, and arms are absent. The spines are particularly well developed in sea urchins. The skeleton, made up of calcium carbonate, may form a large proportion of the body, or, as in some sea cucumbers, it may be greatly reduced. The fossil record shows that the five-rayed pattern is a late evolutionary acquisition, and deviations from it are common. The animals have a well-developed gut, but the nervous system and circulatory system are simple. Echinoderms are slow-moving and without complex behaviour patterns.

III

Ecology

Echinoderms are common on the ocean bottom at all depths; in the deep sea they often make up the bulk of living material. They may be grazers (most sea urchins), feeders on small particles (many brittle stars), or predators (most starfish). Starfish and a few others are pests, as when found in oyster beds. Sea cucumbers and sea urchins are eaten by human beings.

Scientific classification: Echinoderms make up the phylum Echinodermata. Most of the living echinoderms are grouped in the classes: Crinoidea, or sea lilies and feather stars, with about 650 species; Holothuroidea, or sea cucumbers, with about 900 species; Echinoidea, or sea urchins and sand dollars, with about 850 species; Asteroidea, or starfish, with about 1,500 species; and Ophiuroidea, or brittle stars, with about 2,000 species. Many authorities, however, unite Asteroidea and Ophiuroidea into a single class, Asterozoa (or Stelleroidea). About 20 extinct classes are found in the fossil record.

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